M I C H A E L     H A S S L E R     F L O R A

Checklist of Ferns and Lycophytes of the World

In cooperation with Botanical Garden of the Karlsruhe Institute of Technology
Disclaimer: The author of the website does not take over responsibility for the correct determination of species based on the information or photos contained on this website, and any impact resulting from this, e.g. in the consumption of species shown in the picture for medical purposes.
Der Autor übernimmt keine Verantwortung für die korrekte Bestimmung von Arten basierend auf Informationen oder Bildern, die auf dieser Website enthalten sind sowie möglicher Folgen, die sich daraus ergeben, z.B. der medizinischen Verwendung von dargestellten Pflanzen.

Checklist of Ferns and Lycophytes of the World

by Michael Hassler and Bernd Schmitt

Fern statistics by country

 

Status:    ++        complete coverage, modern flora existing

              + (+)     modern checklist existing, but modern flora only partially existing

              +          modern checklist existing

              o          coverage partial

              o (+)     coverage partial, but modern flora partially existing

              -           no good or modern treatment for the country available

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

EUROPE

154

22

 

10.291.830

Europe in general:

Flora Europaea, Vol. 1 (2nd ed.) (Tutin et al., ed., 1993)

Checklist of European Pteridophytes (Derrick et al. 1987)

Albania

46

0

++

28.748

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Austria

68

0

++

83.858

 

Baleares

31

2

++

4.912

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Guia de helechos de la Peninsula Iberica y Baleares (Tierra 1990)

Baltic States (Estonia, Lithuania, Latvia)

49

0

++

45.227 (Est.)

63.700 (Latv.)

65.300 (Lith.)

Vascular plants of Russia and adjacent states (Czerepanov 1995, Checklist)

Flora of Russia. The European Part and Bordering Regions. Vol. 1 (Fedorov 1999)

Belgium

53

0

++

30.528

Nouvelle Flore de la Belgique, du Grand-Duché de Luxembourg, du Nord de la France et des régions voisines (De Langhe et al. 1983)

Bulgaria

58

0

++

110.994

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Corsica

64

0

++

8.680

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Crete

39

1

++

8.335

Exkursionsflora für Kreta (Jahn & Schönfelder 1995)

Med-Checklist, Vol. 1 (Greuter et al. 1984)

former Czechoslovakia (Czech Republik & Slovakia)

63

0

++

78.864 (C.R.)

49.035(Slov.)

 

Denmark

46

0

++

43.094

Den danske Flora (Hansen et al. 1975)

Flora Nordica, Vol. 1 (Jonsell, ed., 2000)

England (including Scotland, excluding Northern Ireland)

66

1

++

244.110

The illustrated field guide to ferns and allied plants of the British Iles (Jermy 1991)

Flora of the British Isles (Clapham et al. 1962)

European Turkey

24

0

++

ca. 40.000

Flora of Turkey and the east Aegaean Islands, Vol. 1 (Davis, ed., 1965)

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Faroer Isl.

21

0

++

1.399

 

Finland

55

0

++

338.144

Flora Nordica, Vol. 1 (Jonsell, ed., 2000)

France (except Corsica)

107

1

++

535.285

Med-Checklist, Vol. 1 (Greuter et al. 1984) (southern France only)

Index synonymique de la flore de France (Kerguélen 1993)

Germany

73

0

++

356.970

 

Greece (including Western Aegaean, except Eastern Aegaean Islands)

60

1

++

123.600

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Hungary

54

0

++

93.030

 

Iceland

 

35

0

++

103.000

A guide to the flowering plants and ferns of Iceland (Kristinsson 1998)

Flora Nordica, Vol. 1 (Jonsell, ed., 2000)

Ireland (incl. Northern Ireland)

49

0

++

70.000

The illustrated field guide to ferns and allied plants of the British Iles (Jermy 1991)

Italy (mainland only)

107

1

++

203.856

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Jan Mayen

3

0

++

380

The flora of Jan Mayen (Lid 1964)

Malta

27

0

++

316

Med-Checklist, Vol. 1 (Greuter et al. 1984)

A Flora of the Maltese Islands (Haslam, Sell & Wolseley 1977)

Netherlands

42

0

++

33.920

 

Norway

64

0

++

385.639

Norsk og svensk flora (Lid 1963)

Flora Nordica, Vol. 1 (Jonsell, ed., 2000)

Poland

64

0

++

312.684

 

Portugal

62

2

++

89.357

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Guia de helechos de la Peninsula Iberica y Baleares (Tierra 1990)

Romania

58

0

++

237.500

 

European Russia

(excluding Caucasus),

Belarus,

Ukraine (excluding Crimea),

Moldavia

76

0

++

ca. 4.300.000

(Europ. Russia)

603.700 (Ukraine)

207.595 (Belarus)

33.700 (Moldavia)

Vascular plants of Russia and adjacent states (Czerepanov 1995, Checklist)

Flora of Russia. The European Part and Bordering Regions. Vol. 1 (Fedorov 1999)

Flora of the Russian Arctic, Vol. 1 (Tolmachev, ed., 1995)

 

Sardinia

45

0

++

23.318

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Sicily

45

0

++

25.426

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Spain

100

1

++

498.717

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Plantas Vasculares de la Peninsula Iberica e Islas Baleares, Vol. 1 (Castroviejo et al. 1986)

Checklist of Vascular Plants of Spain, Vol. 1 (Smythies 1984)

Guia de helechos de la Peninsula Iberica y Baleares (Tierra 1990)

Svalbard (Spitzbergen)

5

0

++

62.160

Flora Nordica, Vol. 1 (Jonsell, ed., 2000)

Svalbards flora (Roenning 1964)

Sweden

61

0

++

449.964

Norsk og svensk flora (Lid 1963)

Flora Nordica, Vol. 1 (Jonsell, ed., 2000)

Switzerland

70

0

++

41.285

Flora Helvetica (Lauber & Wagner 1998)

former Yugoslavia (Slovenia, Croatia, Bosnia-Hercegovina, Serbia, Macedonia)

83

2

++

255.000

Med-Checklist, Vol. 1 (Greuter et al. 1984)

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

MACARONESIA

93

17

 

14.294

Macaronesia in general:

Flora of Macaronesia, Checklist of Vascular Plants, 4th ed. (Hansen & Sunding 1993)

Azores

50

4

++

2.247

Flora Europaea, Vol. 1 (2nd ed.) (Tutin et al. 1993)

Canary Islands

57

2

++

7.273

Exkursionsflora für die Kanarischen Inseln (Hohenester & Weiss 1993)

Madeira (incl. Ilhas Desertas, Porto Santo, Salvajes Isl.)

60

5

++

741

Flora of Madeira (Press 1994)

Capverdes

45

0

++

4.033

Flora of Macaronesia, Checklist of Vascular Plants, 4th ed. (Hansen & Sunding 1993)

The Ferns and Fern-Allies (Pteridophyta) of the Cape Verde Islands (Lobin et al. 1998)

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

NORTHWEST AFRICA

65

1

 

4.756.888

Northwest Africa in general:

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Flore de l'Afrique du Nord, Vol. 1 (Maire 1952)

Flore et végétation du Sahara. 3. ed. (Ozenda 1991)

Algeria

51

0

++

2.381.740

Nouvelle Flore de l'Algerie (Quezel & Santa 1962)

Libya

15

0

++

1.757.000

Flora of Libya - Pteridophytes (1989)

Morocco

53

1

++

453.730

Pteridophyta, in: Flore pratique du Maroc, Vol. 1 (Salvo & Ouyahya 1999)

 

Tunisia

28

0

+

164.418

 

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists of Ferns and Fern Allies

TROPICAL AND NORTHEAST AFRICA

742

373

 

22.833.594

 

Angola

199

2

++

1.246.700

Conspectus florae angolensis: Pteridophyta (Schelpe 1977)

Annobon Isl.

31

0

o

ca. 1.500

 

Ascension Isl.

11

6

++

88

Farne auf atlantischen Inseln (Benl & Benl 1988)

The Endemic Flora of St. Helena (Cronk 2000)

Benin (Dahomey)

37

0

++

112.622

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Flora of Benin, Vol. 1 (Souza 1987)

Les Pteridophytes de l'Afrique Intertropicale Francaise (Tardieu-Blot 1953)

Bioko Isl. (Fernando Poo)

204

3

++

2.017

The ferns and fern-allies of West Tropical Africa (Alston 1959)

The Pteridophyta of Bioko, Vol. 1-5 (Benl 1978-91)

Botswana

38

0

++

581.730

Flora Zambesiaca: Pteridophyta (Schelpe 1970)

Burkina Faso (Upper Volta)

12

1

+

274.200

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Catalogue des plantes vasculaires du Burkina Faso (Lebrun 1991)

Flore et végétation du Sahara. 3. ed. (Ozenda 1991)

Burundi

?100-150

[93 in list]

5

- (+)

27.834

Flore de l'Afrique Centrale (various volumes, see literature list, only a minor part of the Ferns is covered)

Cameroun

278

10

++

475.442

Flore du Cameroun, Vol. 3 (Tardieu-Blot 1964)

Les Pteridophytes de l'Afrique Intertropicale Francaise (Tardieu-Blot 1953)

Central African Republic

112

2

+

622.436

Catalogue de la flore de Centrafrique (Boulevert 1977)

Les Pteridophytes de l'Afrique Intertropicale Francaise (Tardieu-Blot 1953)

Chad

30

1

+

1.284.000

Catalogue des plantes vasculaires du Tchad meridional (Lebrun et al. 1972, with supplements 1976, 1977, 1986)

Flore et végétation du Sahara. 3. ed. (Ozenda 1991)

Les Pteridophytes de l'Afrique Intertropicale Francaise (Tardieu-Blot 1953)

Congo (Republic Congo)

214

11

+

342.000

Catalogue des plantes vasculaires du Congo (Sita & Moutsambote 1988)

Les Pteridophytes de l'Afrique Intertropicale Francaise (Tardieu-Blot 1953)

Djibouti

13

0

+

23.200

Catalogue des plantes vasculaires de la République de Djibouti (Lebrun et al. 1990)

Adumbratio Florae Aethiopicae (Pichi-Sermolli 1954 ff.)

[Flora of Ethiopia and Eritrea, Vol. 1: due to be published in 2002]

Egypt (except Sinai peninsula)

12

0

+

938.915

A Manual Flora of Egypt (Muschler 1912)

Flora of Egypt, Vol. 1 (Boulos 1999)

Conspectus Florae Orientalis, Vol. 9 (Zohary, Heyn & Heller 1994)

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Equatorial Guinea (continental part [former Rio Muni], except Bioko Isl. and Annobon Isl.)

?100-120

[80 in list]

0

-

24.500

 

Eritrea

13

1

o

121.144

Adumbratio Florae Aethiopicae (Pichi-Sermolli 1954 ff.)

[Flora of Ethiopia and Eritrea, Vol. 1: due to be published in 2002]

Ethiopia

134

9

o

1.133.380

Adumbratio Florae Aethiopicae (Pichi-Sermolli 1954 ff.)

[Flora of Ethiopia and Eritrea, Vol. 1: due to be published in 2002]

Gabon

168

1

++

267.667

Flore du Gabon, Vol. 8. Ptéridophytes (Tardieu-Blot 1964)

Les Pteridophytes de l'Afrique Intertropicale Francaise (Tardieu-Blot 1953)

Gambia

17

0

o

11.295

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Ghana

146

2

o

238.500

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Guinea

173

3

o

245.857

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Les Pteridophytes de l'Afrique Intertropicale Francaise (Tardieu-Blot 1953)

Guinea-Bissau

6

0

o

36.125

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Ivory Coast (Cote d'Ivoire)

159

2

o

322.462

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Les Pteridophytes de l'Afrique Intertropicale Francaise (Tardieu-Blot 1953)

Kenya

258

3

+ (+)

582.646

Flora of Tropical East Africa (various volumes)

Pteridophytes of Tropical East Africa (Johns 1991, Checklist)

Liberia

165

2

+

99.067

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Über die Farne Liberias (Westafrika), ihr Vorkommen und ihre Verbreitung (Kunkel 1962)

Malawi

187

0

++

118.484

Flora Zambesiaca: Pteridophyta (Schelpe 1970)

Mali

31

0

+

1.240.192

Catalogue des plantes vasculaires du Mali (Boudet et al. 1986)

Flore et végétation du Sahara. 3. ed. (Ozenda 1991)

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Mauretania

8

0

++

1.030.700

Catalogue des plantes vasculaires de la Mauritanie et du Sahara occidental (Lebrun 1998)

Flore de Mauritanie, Vol. 2 (Barry & Celles 1991)

Flore et végétation du Sahara. 3. ed. (Ozenda 1991)

Mozambique

206

1

++

799.380

Flora Zambesiaca: Pteridophyta (1970)

Niger

14

0

+

1.267.000

Catalogue des plantes vasculaires du Niger (Fabrègues & Lebrun 1976)

Flore et végétation du Sahara. 3. ed. (Ozenda 1991)

Les Pteridophytes de l'Afrique Intertropicale Francaise (Tardieu-Blot 1953)

Nigeria

180

1

o

923.768

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Les Pteridophytes de l'Afrique Intertropicale Francaise (Tardieu-Blot 1953)

Pemba Isl.

 14

0

+ (+)

982

Flora of Tropical East Africa (various volumes)

Pteridophytes of Tropical East Africa (Johns 1991, Checklist)

Principe Isl.

80

3

+

146

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Catalogue of the vascular plants of S. Tome etc. (Exell 1944)

Checklist of the ferns of Sao Tome and Principe (Internet)

Rwanda

?100-150

[79 in list]

3

- (+)

26.338

Flore de l'Afrique Centrale (various volumes, see literature list, only a minor part of the Ferns is covered)

Sao Tome

146

7

+

855

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Catalogue of the vascular plants of S. Tome etc. (Exell 1944)

Checklist of the ferns of Sao Tome and Principe (Internet)

Senegal

40

0

+

196.722

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Enumeration des plantes vasculaires du Senegal (Lebrun 1973)

Les Pteridophytes de l'Afrique Intertropicale Francaise (Tardieu-Blot 1953)

Sierra Leone

130

0

o

71.740

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Somalia

34

0

++

637.700

The plants of Somalia: an overview and checklist (Kuchar 1986)

Flora of Somalia, Vol. 1 (Thulin, ed., 1993)[*1] 

St. Helena

32

13

++

122

The Endemic Flora of St. Helena (Cronk 2000)

St. Helena: Checklist of endemic & indigenous plants (Clifton 1996)

Farne auf atlantischen Inseln (Benl & Benl 1988)

Sudan

129

0

+

2.505.800

The ferns and the fern-allies of the Sudan (Macleay 1953)

Les Pteridophytes de l'Afrique Intertropicale Francaise (Tardieu-Blot 1953)

Tanzania (except islands)

371

15

+ (+)

942.626

Flora of Tropical East Africa (various volumes)

Pteridophytes of Tropical East Africa (Johns 1991, Checklist)

Togo

51

0

+

56.785

The ferns and fern-allies of West Tropical Africa (Alston 1959)

Tristan da Cunha (incl. surrounding islands)

36

11

+

202

The terrestrial botany of the Tristan da Cunha Islands (Wace & Dixon 1965)

Farne auf atlantischen Inseln (Benl & Benl 1988)

Uganda

252

4

+ (+)

241.038

Flora of Tropical East Africa (various volumes, see literature list)

Pteridophytes of Tropical East Africa (Johns 1991, Checklist)

Western Sahara (Spanish Sahara)

0

0

++

267.769

Catalogue des plantes vasculaires de la Mauritanie et du Sahara occidental (Lebrun 1998)

Flore de Mauritanie, Vol. 2 (Barry & Celles 1991)

Flore et végétation du Sahara. 3. ed. (Ozenda 1991)

Democratic Republic of Congo (Zaire)

?250-300

[229 in list]

13

- (+)

2.344.885

Flore de l'Afrique Centrale (various volumes, see literature list) [only a minor part of the Ferns is covered]

Pteridophytes of the Upper Katanga (Kornas et al. 2000)

Zambia

141

3

++

752.614

Flora Zambesiaca: Pteridophyta (1970)

Distribution and ecology of the Pteridophytes in Zambia (Kornas 1978)

Zanzibar

26

0

+ (+)

1.660

Flora of Tropical East Africa (various volumes)

Pteridophytes of Tropical East Africa (Johns 1991, Checklist)

Zimbabwe

238

1

++

390.759

Flora Zambesiaca: Pteridophyta (1970)

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

SOUTHERN AFRICA

261

45

 

2.091.077

Southern Africa in general:

Vascular Plants of Southern Africa: Names and Distribution (Arnold & de Wet 1993)

Flora of Southern Africa: Pteridophyta (Schelpe & Anthony 1986)

South Africa

257

26

++

1.219.090

 

Lesotho

65

0

++

30.355

 

Namibia

53

0

++

824.269

Prodromus einer Flora von Südwestafrika, Lfg. 26 (Merxmüller, ed., 1969)

Swaziland

77

0

++

17.363

 

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

INDIAN OCEAN

696

377

 

601.231

 

Comores

168

7

+ (+)

1.862

Flore de Madagascar et des Comores, pt. 1-11,13 (Tardieu-Blot 1958-71)

Flore de Madagascar et des Comores, pt. 14 (Stefanovic et al. 1997)

Madagascar

579

245

+ (+)

587.041

Flore de Madagascar et des Comores, pt. 1-11,13 (Tardieu-Blot 1958-71)

Flore de Madagascar et des Comores, pt. 14 (Stefanovic et al. 1997)

Mascarenes (Mauritius,

La Réunion, Rodrigues)

283

48

+

4.484

Les Fougères des Mascareignes et des Seychelles (Tardieu-Blot 1960)

The pteridophytes of Reunion Island (Badre & Cadet 1978)

Seychelles

94

10

+

454

Les Fougères des Mascareignes et des Seychelles (Tardieu-Blot 1960)

Chagos Arch. (Diego Garcia)

7

0

+

30

An annotated checklist of the flora of Diego Garcia, British Indian Ocean Territory (Topp 1988)

Marion & Prince Edward Isl.

9

2

+

ca. 100

Biota Australis Terrestris (Ashworth & Vestal, Internet database)

Ile Amsterdam, St. Paul Isl.

10

1

+

67

Biota Australis Terrestris (Ashworth & Vestal, Internet database)

Iles Crozet

6

0

+

300

Biota Australis Terrestris (Ashworth & Vestal, Internet database)

Iles Kerguelen

8

1

+

6.993

Biota Australis Terrestris (Ashworth & Vestal, Internet database)

Heard & McDonald Isl.

0

0

+

ca. 100

Biota Australis Terrestris (Ashworth & Vestal, Internet database)

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

ARABIAN PENINSULA & SOCOTRA

54

1

 

3.194.603

Arabia in general:

Flora of the Arabian Peninsula and Socotra, Vol. 1 (Miller & Cope 1996)

Conspectus Florae Orientalis, Vol. 9 (Zohary, Heyn & Heller 1994)

Kuwait

1

0

++

17.820

 

Bahrain

2

0

++

707

 

Oman (Mascat, Oman, Dhofar)

15

0

++

309.500

 

Qatar

0

0

++

11.427

 

Saudi Arabia

28

0

++

2.240.000

Flora of Eastern Saudi-Arabia (Mandaville 1990)

Flora of Saudi-Arabia, Vol. 1 (Migahid 1988)

Socotra (incl. Abd-al-Kuri Isl.)

24

1

++

3.579

 

United Arab Emirates

2

0

++

83.600

 

Yemen

44

0

++

527.970

 

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

WESTERN ASIA

132

8

 

3.606.622

 

Afghanistan

33

0

-

652.225

 

Asiatic Turkey (Anatolia)

98

2

++

ca. 74.000

Flora of Turkey and the east Aegaean Islands, Vol. 1 (Davis, ed., 1965)

Flora of Turkey and the east Aegaean Islands, Vol. 10 (Suppl. 1) (Davis, ed., 1988)

Flora of Turkey and the east Aegaean Islands, Vol. 11 (Suppl. 2) (Güner et al., ed., 1988)

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Conspectus Florae Orientalis, Vol. 9 (Zohary, Heyn & Heller 1994)

Caucasus and Transcaucasus (Georgia, Armenia, Aserbaidshan, Talysh region, Caucasian Republics of Russia [Daghestan, Chechnya etc.])

86

3

+

69.700 (Ge.)

29.800 (Arm.)

86.600 (As.)

200.000 (Ca.)

Vascular plants of Russia and adjacent states (Czerepanov 1995, Checklist)

Flora Armenii, Vol. 1 (Tahtadzjana, ed., 1954)

 

Crimea peninsula

40

0

++

25.993

Flora of Russia. The European Part and Bordering Regions. Vol. 1 (Fedorov 1999)

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Vascular plants of Russia and adjacent states (Czerepanov 1995, Checklist)

Cyprus

25

0

++

9.251

Flora of Cyprus, Vol. 1 (Meikle 1985)

Med-Checklist, Vol. 1 (Greuter et al. 1984)

An Illustrated Flora of North Cyprus, Vol. 2 (Viney 1996)

Conspectus Florae Orientalis, Vol. 9 (Zohary, Heyn & Heller 1994)

Eastern Aegaean Islands (Rhodos, Karpathos, Kos, Samos, Lesbos, Chios, Ikaria, Limnos etc.)

26

0

++

ca. 7.000

Flora of Turkey and the east Aegaean Islands, Vol. 1 (Davis, ed., 1965)

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Iran

56

0

+

1.648.000

Conspectus Florae Orientalis, Vol. 9 (Zohary, Heyn & Heller 1994)

Iraq

19

0

++

438.317

Flora of Iraq, vol. 2 (E.Guest et al., ed., 1966)

Flora of lowland Iraq (Rechinger 1964)

Conspectus Florae Orientalis, Vol. 9 (Zohary, Heyn & Heller 1994)

Israel and Palestine

20

0

+

21.946

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Conspectus Florae Orientalis, Vol. 9 (Zohary, Heyn & Heller 1994)

Jordania (except West Bank)

5

0

+

89.556

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Conspectus Florae Orientalis, Vol. 9 (Zohary, Heyn & Heller 1994)

Lebanon

33

0

+

10.230

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Nouvelle Flore du Liban et de la Syrie I (Mouterde 1966)

Conspectus Florae Orientalis, Vol. 9 (Zohary, Heyn & Heller 1994)

Sinai peninsula

7

0

+

58.824

Med-Checklist, Vol. 1 (Greuter et al. 1984)

A Manual Flora of Egypt (Muschler 1912)

Conspectus Florae Orientalis, Vol. 9 (Zohary, Heyn & Heller 1994)

Syria

18

0

+

185.180

Med-Checklist, Vol. 1 (Greuter et al. 1984)

Nouvelle Flore du Liban et de la Syrie I (Mouterde 1966)

Conspectus Florae Orientalis, Vol. 9 (Zohary, Heyn & Heller 1994)

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

NORTH, CENTRAL

AND EASTERN ASIA

2988

[too high]

1907

 

27.875.469

 

China (incl. Tibet, excl. Taiwan)

2563

[too high]

1405

[too high]

+ (+)

9.571.300

Flora Reipublicae Popularis Sinicae (Vol. 2-6)

Flora of China - Checklist (Missouri Botanical Garden, Internet database)

Manuscript drafts for the upcoming Flora of China (Missouri Botanical Garden, Internet website for Flora of China)

Plants of Central Asia, Vol. 1 (Grubov 1963 [transl. 1999])

Japan (incl. Yakushima Isl. and Izu Isl., except Ryukyu and Bonin Isl.)

721

129

++

372.769

Flora of Japan (Ohwi 1965)

Flora of Japan, Vol. 1 (Iwatsuki et al., ed., 1995)

North Korea, South Korea

241

21

-

120.500

(North Korea)

99.300

(South Korea)

 

Mongolia

48

0

++

1.566.500

Key to the Vascular Plants of Mongolia (Grubov 1982)

Plants of Central Asia, Vol. 1 (Grubov 1963 [transl. 1999])

Ryukyu Isl.

268

14

++

4.700

Flora of the Ryukyu Islands (Walker 1976)

Siberia

(incl. Kamchatka, Kuril Isl., Amur, Ussuri, Sakhalin, Primorye, Magadan, etc.)

193

6

+

ca. 11.500.000 (Siberia)

76.400 (Sakhalin)

15.600

(Kuril Isl.)

518.000 (Kamchatka)

Vascular plants of Russia and adjacent states (Czerepanov 1995, Checklist)

Flora of the Russian Arctic, Vol. 1 (Tolmachev, ed., 1995)

Plants of Central Asia, Vol. 1 (Grubov 1963 [transl. 1999])

Taiwan (Formosa)

738

125

++

36.000

Flora Reipublicae Popularis Sinicae (Vol. 1-6, 1990 ff.)

Flora of Taiwan, Vol. 1 (2nd. ed., 1994)

Central Asian Republics (Kazakstan, Tadjikistan, Turkmenistan, Uzbekistan, Kirgisia)

56

1

++

2.717.300 (Kazakstan)

143.100 (Tadjikistan)

488.100 (Turkmen.)

447.400 (Uzbekistan)

198.500 (Kirgisia)

Vascular plants of Russia and adjacent states (Czerepanov 1995, Checklist)

Plants of Central Asia, Vol. 1 (Grubov 1963 [transl. 1999])

 

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

INDIAN SUBCONTINENT AND INDOCHINA

1598

344

 

6.437.509

 

Andaman Isl.

68

2

++

6.496

Pteridophytes of Andaman and Nicobar Islands (Dixit 2001)

Bangladesh

?100-200

[68 in list]

0

-

147.570

 

Bhutan

258

1

o

47.000

New Species Syndrome in Indian Pteridology and the ferns of Nepal (Fraser-Jenkins 1997)

Burma (Myanmar)

?500-600

[493 in list]

12

-

676.552

 

Cambodia

?300-400

[140 in list]

0

o

181.035

Fougeres (de la Flore Generale de l'Indochine) (Tardieu-Blot & Christensen 1939-1951)

India

1042

93

+ (+)

3.278.967

[over 20 regional floras of India not listed here, see literature list. See also extensive literature list in Khullar 1994/2000]

Illustrated Manual of Ferns of Assam (Borthakur et al. 2000)

A census of the Indian Pteridophytes (Dixit 1984)

The Ferns of India: Enumeration, Synonyms and Distribution (Chandra 2000)

New Species Syndrome in Indian Pteridology and the ferns of Nepal (Fraser-Jenkins 1997)

Fern flora of South India (Nampy & Madhusoodanan 1998)

An illustrated fern flora of West Himalaya. Vol. 1-2 (Khullar 1994/2000)

Laos

?300-400 [140 in list]

1

o

236.800

Fougeres (de la Flore Generale de l'Indochine) (Tardieu-Blot & Christensen 1939-1951)

Maledives

?0

0

-

298

 

Nepal

393

2

+

147.181

An Enumeration of the Flowering Plants of Nepal, Vol. 1 (Hara et al. 1978)

An enumeration of the Pteridophytes of Nepal (Iwatsuki 1988)

New Species Syndrome in Indian Pteridology and the ferns of Nepal (Fraser-Jenkins 1997)

An illustrated fern flora of West Himalaya. Vol. 1-2 (Khullar 1994/2000)

Nicobar Isl.

87

2

++

1.800

Pteridophytes of Andaman and Nicobar Islands (Dixit 2001)

Flora of Great Nicobar Island (Sinha 1999)

Pakistan

138

1

+

796.095

An illustrated fern flora of West Himalaya. Vol. 1-2 (Khullar 1994/2000)

Flora of West Pakistan. Incl.: An annotated catalogue of the vascular plants of West Pakistan and Kashmir (by R. R. Stewart) (Nasir, ed., 1972)

Sikkim

297

3

+ (+)

7.300

A census of the Indian Pteridophytes (Dixit 1984)

The Ferns of India: Enumeration, Synonyms and Distribution (Chandra 2000)

Sri Lanka

396

37

+

65.610

A check list of the pteridophytes of Sri Lanka (Abeywickrama 1978)

An annotated check-list of the Pteridophytes of Ceylon (Sledge 1982)

Thailand

652

21

++

513.115

Flora of Thailand, Vol. 3/1-4 (Tagawa & Iwatsuki 1979-1989)

Pteridophytes in Thailand (Boonkeerd & Pollawatn 2000)

Vietnam

591

66

o

331.690

Fougeres (de la Flore Generale de l'Indochine) (Tardieu-Blot & Christensen 1939-1951)

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

MALESIA

3227

2326

 

2.959.296

Malesia in general (except Bismarck Arch., Solomons, Bougainville):

Flora Malesiana, Series II: Fern and Fern Allies. Vol. 1-3 (to be continued)

Bismarck Arch. (New Britain, New Ireland, New Hanover, Admirality Isl., etc.)

?300-400

[84 in list]

19

- (+)

48.500

 

Borneo (Sarawak, Sabah, Kalimantan, Brunei, incl. Natuna Isl., Riouw Arch.)

1185

722

- (+)

743.384

The Plants of Mt. Kinabalu, vol. 1 (Ferns and Fern Allies) (Parris et al. 1992)

Bougainville Isl.

?100-150

[22 in list]

4

-

8.370

 

Java (incl. Madura Isl.)

519

75

- (+)

130.520

 

Lesser Sunda Isl. (Bali, Lombok, Sumbawa, Flores, Sumba, Timor, Wetar)

164

14

- (+)

ca. 80.000

 

peninsular Malaysia, Singapore (incl. Langkawi Isl.)

660

49

++

132.246

A revised Flora of Malaya II: Ferns of Malaya, ed. 2 (Holttum 1966)

A Guide to the Ferns of Singapore (Wee 1987)

Moluccas (Tanimbar, Kei, Aru, Banda, Ambon, Ceram, Buru, Sula, Obi, Bacan, Halmaheira, Ternate, Tidore, Morotai, Talaud)

?500-600 [352 in list]

64

- (+)

74.500

 

New Guinea (Irian Jaya and Papua New Guinea, incl. d'Entrecasteaux Isl., Louisiade Arch., Rennell Isl.)

1183

722

- (+)

800.000

 

Philippines

1048

402

o (+)

300.000

Guide to Philippine Flora and Fauna, Vol. 2 (Zamora & Co 1986)

Solomons

?300-400

[159 in list]

39

-

27.556

 

Sulawesi (Celebes) (incl. Selayar, Banggai, Sangihe)

?500-700

[397 in list]

82

- (+)

189.070

 

Sumatra (incl. Mentawai, Enggano, Nias, Simeuleu, Lingga, Anambas, Krakatau, Bangka, Belitung)

?700-800

[615 in list]

155

- (+)

425.150

 

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

AUSTRALIA

470

159

 

7.690.849

Australia in general:

Census of Australian Vascular Plants (Hnatiuk 1990)

Flora of Australia, Volume 48 (1998)

New South Wales

177

3

++

801.500

 

Australian Capital Territory

22

0

++

 

 

Northern Territory

84

3

++

1.346.200

 

Queensland

390

74

++

1.727.200

 

South Australia

53

1

++

984.400

Flora of South Australia (Jessop & Toelken 1986)

Ferns and Allied Plants of Victoria, Tasmania and South Australia (Duncan & Isaac 1994)

Tasmania

102

7

++

68.300

Ferns and Allied Plants of Victoria, Tasmania and South Australia (Duncan & Isaac 1994)

The Ferns of Tasmania (Garrett 1997)

Victoria

122

1

++

227.600

Ferns and Allied Plants of Victoria, Tasmania and South Australia (Duncan & Isaac 1994)

Flora of Victoria, Vol. 2 (Walsh & Entwisle, ed., 1994)

Western Australia

85

8

++

2.535.500

 

Christmas Island (Australia)

31

2

++

135

Flora of Australia, Vol. 49/50 (Oceanic Islands) (1993/1994)

Cocos Island (Keeling Island)

0

0

++

14

Flora of Australia, Vol. 49/50 (Oceanic Islands) (1993/1994)

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

PACIFIC

1209

745

 

349.946

Pacific in general:

Vegetation of the Tropical Pacific Islands (Mueller-Dombois & Fosberg 1998)

American Samoa

110

2

o

197

A revision of the Pteridophyta of Samoa (Christensen 1943)

Vegetation of Eastern Samoa (Whistler 1980)

Plants of the Manua Islands (Yuncker 1945)

Antipodes Isl.

23

1

++

ca. 60

New Zealand Ferns and Allied Plants (Brownsey & Smith-Dodsworth 1989)

A revised classification of New Zealand pteridophytes with a synonymic checklist of species (Brownsey, Given & Lovis 1985)

Biota Australis Terrestris (Ashworth & Vestal, Internet database)

Auckland Isl.

43

0

++

551

New Zealand Ferns and Allied Plants (Brownsey & Smith-Dodsworth 1989)

Biota Australis Terrestris (Ashworth & Vestal, Internet database)

A revised classification of New Zealand pteridophytes with a synonymic checklist of species (Brownsey, Given & Lovis 1985)

Austral Isl. (Tubuai Isl., incl. Rapa Isl.)

103

25

+

148

Les Orchidees du Tubuai, suives d'un catalogue des plantes a fleurs et fougeres des Iles Australes (Hallé 1980)

Flora of Southestern Polynesia, II. Pteridophytes (Brown & Brown 1931)

Banaba Isl. (Ocean Isl.)

1

0

+

ca. 10

Geographical checklist of the Micronesian Pteridophyta and Gymnospermae (Fosberg et al. 1982)

Bonin Isl. (incl. Volcano Isl. = Kazan-Retto = Iwo-jima)

79

22

++

132

A revised list of vascular plants indigenous and introduced to the Bonin (Ogasawara) and the Volcano (Kazan) Islands (Kobayashi & Ono 1987)

Flora of the Bonin Islands (Toyoda 1981)

Campbell Isl.

22

0

++

137

New Zealand Ferns and Allied Plants (Brownsey & Smith-Dodsworth 1989)

A revised classification of New Zealand pteridophytes with a synonymic checklist of species (Brownsey, Given & Lovis 1985)

Biota Australis Terrestris (Ashworth & Vestal, Internet database)

Chatham Isl., Snares Isl.

84

1

++

963

New Zealand Ferns and Allied Plants (Brownsey & Smith-Dodsworth 1989)

A revised classification of New Zealand pteridophytes with a synonymic checklist of species (Brownsey, Given & Lovis 1985)

Christmas Isl. (Kiribati p. p.)

0

0

+

321

Plants of Christmas Island (Chock & Hamilton 1962)

Cook Isl. (incl. Rarotonga Isl.)

58

6

+

230

Flora of Rarotonga (Wilder 1931)

Federal States of Micronesia (Central and Eastern Carolines: Truk, Yap, Pohnpei, Kosrae)

134

15

+

702

Geographical checklist of the Micronesian Pteridophyta and Gymnospermae (Fosberg et al. 1982)

Fiji

298

75

++

18.376

The pteridophyte flora of Fiji (Brownlie 1977)

Gambier Isl. (incl. Mangareva Isl.)

9

0

+

ca. 50

Census of the flora of the Gambier Islands, Polynesia (St. John 1988)

Flora of Southestern Polynesia, II. Pteridophytes (Brown & Brown 1931)

Gilbert Islands (Kiribati p. p.)

4

0

+

311

Flora of the Gilbert Islands, Kiribati, checklist (Fosberg & Sachet 1987)

Geographical checklist of the Micronesian Pteridophyta and Gymnospermae (Fosberg et al. 1982)

Vegetation of Nauru and the Gilbert Islands (Thaman 1992)

Hawaii

167

116

+

16.729

Ferns of Hawai’i (Valier 1995)

Henderson Isl. (incl. Ducie Atoll)

8

0

+

37

Henderson Island (Southeast Polynesia) (Fosberg et al. 1983)

Flora of Southestern Polynesia, II. Pteridophytes (Brown & Brown 1931)

Kermadec Isl.

36

3

++

36

Kermadec Islands Flora. An annotated checklist (Sykes 1977)

New records and other information on the vascular flora of the Kermadec Islands (Sykes & West 1996)

A revised classification of New Zealand pteridophytes with a synonymic checklist of species (Brownsey, Given & Lovis 1985)

New Zealand Ferns and Allied Plants (Brownsey & Smith-Dodsworth 1989)

Line Islands (Kiribati p. p.)

5

0

+

436

Notes on the vascular flora and terrestrial vertebrates of Caroline Atoll, Southern Line Islands (Clapp & Sibley 1971)

Vegetation History of Washington Island (Teraina), Northern Line Islands (Wester et al. 1992)

Checklist of the vascular plants of the northern Line Islands (Wester 1985)

Lord Howe Isl.

61

23

++

154

Flora of Australia, Vol. 49/50 (Oceanic Islands) (1993/1994)

Macquarie Isl.

6

0

++

123

Flora of Australia, Vol. 49/50 (Oceanic Islands) (1993/1994)

Biota Australis Terrestris (Ashworth & Vestal, Internet database)

Marcus Isl.

0

0

+

ca. 10

Geographical checklist of the Micronesian Pteridophyta and Gymnospermae (Fosberg et al. 1982)

Marquesas Isl.

ca. 120

[74 in list]

21

o

1.020

Flora of Southestern Polynesia, II. Pteridophytes (Brown & Brown 1931)

Marshall Isl.

10

0

+

174

Geographical checklist of the Micronesian Pteridophyta and Gymnospermae (Fosberg et al. 1982)

Vascular Plants recorded from Jaluit Atoll (Fosberg & Sachet 1962)

A review of the natural history of the Marshall Islands (Fosberg 1990)

Nauru

3

0

++

21

Geographical checklist of the Micronesian Pteridophyta and Gymnospermae (Fosberg et al. 1982)

Vegetation of Nauru and the Gilbert Islands (Thaman 1992)

The Flora of Nauru (Thaman et al. 1994)

New Caledonia (incl. Ile des Pins, Loyalty Isl., Ile Art)

263

107

++

19.058

Flore de la Nouvelle-Caledonie et Dependances, Vol. 3 (Brownlie 1969)

New Zealand (incl. Stewart Isl., Three Kings Isl.)

181

36

++

270.000

New Zealand Ferns and Allied Plants (Brownsey & Smith-Dodsworth 1989)

A revised classification of New Zealand pteridophytes with a synonymic checklist of species (Brownsey, Given & Lovis 1985)

Niue Isl.

28

0

++

264

The flora of Niue Island (Yuncker 1943)

Contributions to the flora of Niue (Sykes 1970)

Norfolk Isl.

47

7

++

39

Flora of Australia, Vol. 49/50 (Oceanic Islands) (1993/1994)

Northern Marianas

28

1

++

152

Geographical checklist of the Micronesian Pteridophyta and Gymnospermae (Fosberg et al. 1982)

Ferns and Orchids of the Mariana Islands (Raulerson & Rinehart 1992)

Palau Isl. (Republic of Palau, Western Carolines)

124

13

+

516

Geographical checklist of the Micronesian Pteridophyta and Gymnospermae (Fosberg et al. 1982)

Vascular Plants of Palau with vernacular names (Fosberg et al. 1980)

Phoenix Islands (Kiribati p.p.)

0

0

++

42

Flora of the Phoenix Islands, Central Pacific (Fosberg & Stoddart 1994)

Pitcairn Isl., Oeno Atoll

18

4

+

6

An account of the flora of Pitcairn Island (St. John 1987)

Flora of Southestern Polynesia, II. Pteridophytes (Brown & Brown 1931)

Society Isl. (Tahiti, Moorea, Raiatea, Bora Bora, Makatea, etc.)

190

67

o

1.628

Pteridophytes of Society Islands (Copeland 1931)

 

Southern Marianas (incl. Guam)

67

4

++

867

Geographical checklist of the Micronesian Pteridophyta and Gymnospermae (Fosberg et al. 1982)

Ferns and Orchids of the Mariana Islands (Raulerson & Rinehart 1992)

The Flora of Guam (Stone 1970)

Tokelau Isl. (incl. Swains Isl.)

9

0

o

14

The vegetation of the Tokelau Islands with special reference to the plants of Nukunonu Atoll (Parham 1971)

The Flora and Vegetation of Swains Island (Whistler 1983)

Tonga

90

3

+

750

Plants of Tonga (Yuncker 1959)

The vegetation of Late, Tonga (Sykes 1981)

The Pteridophytes of 'Eua, Southern Tonga (Sykes 1978)

New or noteworthy ferns from the Tongan Islands (Iwatsuki 1963)

Tuamotu Arch. (excl. Gambier Isl.)

9

0

o

ca. 450

Takapoto Atoll, Tuamotu Archipelago: terrestrial vegetation (Sachet 1983)

Flora of Makatea (Wilder 1934)

Flora of Southestern Polynesia, II. Pteridophytes (Brown & Brown 1931)

Tuvalu (Ellice Isl.)

7

0

o

26

Vegetation and Flora of Nui Atoll, Tuvalu (Woodroffe 1985)

Vanuatu (incl. Santa Cruz Isl.)

?130-170

[120 in list]

22

-

12.190

 

Wake Isl.

0

0

+

8

Vegetation and flora of Wake Island (Fosberg 1959)

Wake Island vegetation and flora, 1961-1963 (Fosberg & Sachet 1969)

Geographical checklist of the Micronesian Pteridophyta and Gymnospermae (Fosberg et al. 1982)

Wallis Isl., Futuna Isl. (Horne Isl.)

59

[6 in list]

 

?0

-

177

 

Western Samoa

217

29

o

2.831

A revision of the Pteridophyta of Samoa (Christensen 1943)

Vegetation and Flora of the Aleipata Islands, Western Samoa (Whistler 1983)

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

NORTH AMERICA

435

138

 

21.938.931

North America in general:

Flora of North America, North of Mexico, Vol. 2 (1993)

A Field Manual of the Ferns and Fern-Allies of the United States and Canada (Lellinger 1985)

A Synonymized Checklist of the Vascular Plants of the United States, Canada and Greenland (Kartesz & Kartesz 1980)

Canada

167

2

++

9.970.610

The flora of Canada (Scoggan 1978)

Les fougeres et les plantes alliees du Canada (Cody & Britton 1989)

USA

410

74

++

8.262.426

 

Alaska (incl. Aleutes)

66

0

++

1.530.000

Flora of Alaska and neighboring territories (Hultén 1968)

Greenland

31

0

++

2.175.600

 

Bermuda

19

4

o

53

Flora of Bermuda (Britton 1918)

St. Pierre & Miquelon Isl.

32

0

++

242

 

 

Country / Region

Species count

endem.

Status

Area (km2)

Reference works and checklists

MEXICO AND CENTRAL AMERICA

2620

616

 

2.485.930

Central America in general:

Flora Mesoamericana, Vol. 1 (1995) [covers in Mexico only Chiapas fully]

Ferns and allied plants, with special reference to tropical America (Tryon & Tryon 1982)

Belize

250

2

++

22.965